Yes I had already tried changing theme and deactivating all plugins before I posted.
But.. I see you’ve reproduced. 🙂
I also noticed (after I downgraded to previous version) that even with registration disabled I was able to load register page. Then I upgraded BP again and it was still a case.
Weird! That does work! Thanks Rhoward!! I switched settings to “Both sites and user accounts can be registered.” and it will save the settings. But if I change back to “Logged in users may register new sites.” the settings are no longer saved. :/
WP Version 4.1
Multisite
It is network activated on the main site
I have the invite anyone plugin installed so it is set to “Logged in users may register new sites.”
WP Super Cache
It is the weirdest thing because it was just working the other day… :/