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Yes, there was a filter on pre_get_posts, from another plugin:/

This can be closed!

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UPDATE: well, I found the problem. It seems to be a bug(?) in bp_get_email. That function should get the email-template from the main/root blog. When another blog is active, it switches to the rootblog for the template-query. That switch (switch_to_blog) doesnt seem to work.. So it only works on the rootblog, not on the other blogs (multisite).

Next I checked the difference between the queries on the rootblog and the other blogs. The difference is in the Post_type statement:
– On rootblog: wp_posts.post_type IN (‘bp-email’)
– On other blogs: wp_posts.post_type IN (‘post’)

The bp_get_email_post_type and bp_get_email_tax_type do return the right values when switched to the rootblog, but maybe there’s some filter on WP_Query where the posttypes are verified or something?

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Yep, that option is selected. I think an error occures somewhere along, but its hard to debug this flow. Any ideas are welcome…

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@pixelpushermama @julia_B did you fix this, or is it still a problem?

I just upgraded from BuddyPress 2.3.4 to 2.7.4 (WP 4.7.1) but now we seem to have a problem with the mentions. Users used to get an notification-email when they were mentioned in the comments on one of our sites. That doesnt happen anymore. But the emails from private messages or replies on activity-updates do work…