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Solved. I’m an idiot.
user_spam_remover Plugin was improperly configured.
As a side note: If you use buddypress for commenting, the User Spam Remover plugin won’t pick up on buddypress activity as being actual site activity – thus causing users to be mistaken as spam bots.
See also: WordPress.org • bbPress.org • BuddyPress.org • Matt • Blog RSS