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Here we go – easier than I thought it would be to fix this.
Go to bp-core/bp-core-signup.php and find the following line:
$account_details = bp_core_validate_user_signup( $_POST['signup_username'], $_POST['signup_email'] );
Add this line directly below it:
$account_details=strtolower($account_details);
It just transforms usernames typed in multiple cases in the username field to only lowercase letters.